When this item is selected (displays a check mark in the menu), Finale assumes that you'll be entering notes in conjunction with a MIDI keyboard. When you press one of the number (rhythmic value) keys on the numeric keypad, Finale creates a rest of that value, unless you were pressing a key on the synthesizer at the time. In that case, Finale displays a note of that pitch (or a chord, if you were pressing more than one key).
When Use MIDI Keyboard isn't selected, Finale enters a note of the specified rhythmic value when you press a number key - regardless of whether or not you were pressing a key on the synthesizer (if any). To create a rest, enter a single note of the desired rhythmic value and press BACKSPACE.
See also:
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